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© 2017 Patrick Bateson

The text of this book is licensed under a Creative Commons Attribution-NonCommercial-NoDerivatives 4.0 International license (CC BY-NC-ND 4.0). This license allows you to share, copy, distribute and transmit the work for non-commercial purposes, providing attribution is made to the author (but not in any way that suggests that he endorses you or your use of the work). Attribution should include the following information:

Bateson, Patrick, Behaviour, Development and Evolution. Cambridge, UK: Open Book Publishers, 2017.

In order to access detailed and updated information on the license, please visit

Further details about CC BY-NC-ND licenses are available at

All external links were active on 20/2/2017 unless otherwise stated and have been archived via the Internet Archive Wayback Machine at

Updated digital material and resources associated with this volume are available at

Every effort has been made to identify and contact copyright holders and any omission or error will be corrected if notification is made to the publisher.

ISBN Paperback: 978-1-78374-248-6

ISBN Hardback: 978-1-78374-249-3

ISBN Digital (PDF): 978-1-78374-250-9

ISBN Digital ebook (epub): 978-1-78374-251-6

ISBN Digital ebook (mobi): 978-1-78374-252-3

DOI: 10.11647/OBP.0097

Cover image: Johannes Itten, ‘Cerchi’ (1916) © Galleria Nazionale d’Arte Moderna e Contemporanea, Rome, all rights reserved. Su concessione del Ministero dei Beni e delle Attività Culturali e Ambientali e del Turismo. Photo by Anna Gatti.

All paper used by Open Book Publishers is SFI (Sustainable Forestry Initiative), and PEFC (Programme for the endorsement of Forest Certification Schemes) Certified.

Printed in the United Kingdom, United States and Australia by Lightning Source for Open Book Publishers (Cambridge, UK).